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Other scenarios I can think of as possibilities:
-Replying user doesn't have Append/Append To privileges
-Field is cleared by a Business Rule
-Maybe some kind of caching issue (would recommend trying in a private browser)
You saying that you moved to Dynamics Online from old version of Dynamic CRM or may be On-premise.
Try this for troubleshooting
If regarding is having no problem with new emails, I suspect something went wrong when doing migration.
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Finally figured out the problem. I had a chrome extension that was causing the problem. When I disabled regardingobjectid was there :)
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