Browsing the web application is fast for an end user e.g. when visiting http://applicationurl/orgname/main.aspx# and accessing lists of records, forms, views etc. all load within a few seconds. Any form of customization e.g. when accessing http://applicationurl/orgname/tools/solution/edit.aspx page load times increase to 1-2 minutes. CPU and memory usage on the web server seem normal (< 10% and 50% respectively) likewise for the separate SQL server.
Monitoring network traffic in Chrome shows a long wait period after the initial request:

(there is another lengthy wait for the second page request (areas.aspx?...).
It is a relatively new on-premise instance with two organisations (the second is unused with only the activity feeds solution installed, i.e. does not include a managed solution like the first). Both organisations suffer from the same poor performance when attempting to customize.
Any ideas would be greatly appreciated.
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